Substrate licking might mediate gustation rather than vomeronasal function, particularly in iguanian lizards. It is, therefore, premature to assume the latter. Insufficient data exist to distinguish between taste and vomeronasal function as the basis for chemosensory-mediated behavior in lizards. They occur most frequently on the tongue tip and the ventrolateral margins of the foretongue, though they may be found anywhere. Lingual taste buds are more or less restricted to regions of thick, stratified squamous epithelium. These observations are contrary to statements in the literature which have assumed taste buds to be rare or absent in lizards. Taste buds may be abundant, particularly in the Iguanidae, in which densities greater than 104/ mm2 occur. It is true that the tip and edges of the tongue are particularly sensitive to tastes, as these areas contain many tiny sensory organs called taste buds. They are found on the tongues of all remaining species except Gonatodes antillensis (Gekkonidae) and in the oral epithelia of all species except Chamaeleo jacksoni (Chamaeleonidae). Taste buds occur in all species examined except Varanus indicus (Varanidae). The tongue and oral epithelium beneath and lateral to the tongue have been examined in 37 species of lizard representing all families except the Helodermatidae and Lanthonotidae.
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